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Leo has an inner desire to be special and important, to shine and be the center of attention, to direct or lead rather than follow. Leo wants to stand out from the crowd in some way, and is generally not a team player. Aquarius, on the other hand, needs to be part of a group or community. The fellowship, the togetherness, and the group solidarity are very satisfying and meaningful to Aquarius, whether or not Aquarius is the focus of attention.
Leo wants to be in control and can be very authoritarian (in a magnanimous sort of way, as in a "benign dictatorship"), but Aquarius has a rather rebellious, freedom-loving nature and resists Authority. Though having no desire to be on top, Aquarius will not be compliant or domesticated either.
Also, Leo is loyal, intensely personal, and rather narcissistic, while Aquarius can be rather cool and impartially fair. Leo sometimes feels hurt by being treated "just like everyone else"!